+Present address: School of Biosciences, University of Birmingham, Edgbaston, Birmingham B15 2TT, England.
++Present address: Sareum Ltd, 2 Iconix Park, Pampisford, Cambridge CB22 3EG, England.
+++Present address: Faculty of Biosciences and Engineering, Universiti Teknologi Malaysia, 81310 Skudai, Johor, Malaysia.